Did Neanderthals “become” Rh negative?

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Neanderthal, (Homo neanderthalensis, Homo sapiens neanderthalensis), also spelled Neandertal, member of a group of archaic humans who emerged at least 200,000 years ago during the Pleistocene Epoch (about 2.6 million to 11,700 years ago) and were replaced or assimilated by early modern human populations (Homo sapiens)

This was suggested by A.E. Mourant:

We were interested in discovering where Rh-negative may have originated. We therefore drew up a “weather-map” of the genic distribution of Rhnegative. The resulting isogenic map led us to conclude that Rh-negative originated in Southwestern France. We conjectured the period at about 50,000 years, and the population probably a Neanderthaloid-like (!) one.

Here is the study:

So far, all of the Neanderthal specimens examined were found to be Partial D:

Did the Neanderthals evolve into Rh negatives in at least one region?

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