This was suggested by A.E. Mourant:
We were interested in discovering where Rh-negative may have originated. We therefore drew up a “weather-map” of the genic distribution of Rhnegative. The resulting isogenic map led us to conclude that Rh-negative originated in Southwestern France. We conjectured the period at about 50,000 years, and the population probably a Neanderthaloid-like (!) one.
Here is the study:
So far, all of the Neanderthal specimens examined were found to be Partial D:
Did the Neanderthals evolve into Rh negatives in at least one region?
About the author: